Browsing by Subject "male genitalia"
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Item The caddisfly genus Protoptila in Costa Rica (Trichoptera: Glossosomatidae)(Magnolia Press, 2006) Holzenthal, Ralph W.; Blahnik, Roger J.Nineteen species of Protoptila known from Costa Rica are revised to include 11 previously described species (P. bicornuta Flint 1963, P. boruca Flint 1974, P. burica Flint 1974, P. cana Flint 1974, P. ixtala Mosely 1937, P. laterospina Flint 1967, P. orotina orotina Flint 1974, P. spirifera Flint 1974, P. talamanca Flint 1974, P. tica Bueno-Soria 1984, and P. tojana Mosely 1954) and 8 new species (P. altura, P. bribri, P. chitaria, P. cristula, P. kjeri, P. jolandae, P. strepsicera, and P. trichoglossa). Illustrations, diagnoses, descriptions, and distribution records are provided for all species.Item The Neotropical caddisfly genus Canoptila (Trichoptera: Glossosomatidae)(Magnolia Press, 2006) Robertson, Desiree R.; Holzenthal, Ralph W.The caddisfly genus Canoptila Mosely (Glossosomatidae: Protoptilinae), endemic to southeastern Brazil, is diagnosed and discussed in the context of other protoptiline genera, and a brief summary of its taxonomic history is provided. A new species, Canoptila williami, is described and illustrated, including a female, the first known for the genus. Additionally, the type species, Canoptila bifida Mosely, is redescribed and illustrated. There are three possible synapomorphies supporting the monophyly of Canoptila: 1) the presence of long spine-like posterolateral processes on tergum X; 2) the highly membranous digitate parameres on the endotheca; and 3) the unique combination of both forewing and hind wing venational characters.Item The Neotropical caddisfly genus Tolhuaca (Trichoptera: Glossosomatidae)(Magnolia Press, 2005) Robertson, Desiree R.; Holzenthal, Ralph W.The caddisfly genus Tolhauca Schmid 1964 (Glossosomatidae: Protoptilinae) is diagnosed and discussed in the context of other protoptiline genera, and a review of its taxonomic history is provided. A new species, Tolhuaca brasiliensis, from southeastern Brazil, is described and illustrated, and the type species, Tolhuaca cupulifera Schmid 1964, from Chile, is redescribed and illustrated. Additionally, females of the genus are described and illustrated for the first time. Characters of the female genitalia, wing venation, and thorax suggest that Tolhuaca is more primitive than any other protoptiline genus and probably deserves a basal placement within the subfamily. The genus shows a broadly disjunct distribution perhaps reflecting an ancient southern Gondwana pattern.Item Review of the caddisfly genus Tagalopsyche with the description of new species and a related new genus (Trichoptera: Leptoceridae: Mystacidini)(Magnolia Press, 2007) Holzenthal, Ralph W.; Andersen, TrondThe long-horned caddisfly genus Tagalopsyche Banks, 1913 (Leptoceridae: Mystacidini) is reviewed and 4 new species are described and illustrated: T. apratita, new species (India), T. jolandae, new species (Brunei, Malaysia), T. kjaerandseni, new species (Ghana), and T. udagama, new species (India, Sri Lanka). The 4 previously described species are redescribed and illustrated: T. aethiopica Kimmins, 1963a (Ethiopia), T. brunnea (Ulmer, 1905) (Indonesia), T. fletcheri Kimmins, 1963b (India), and T. sisyroides Banks, 1913 (Indonesia, Philippines). A related new genus, Fernandoschmidia, new genus, and 2 new species, F. amudita, new species, and F. aramaniya, new species, from India are described and illustrated. The phylogenetic position of the new genus is assessed; it is sister to a clade formed by Tagalopsyche and Mystacides Berthold, 1827, the only other genus in the Mystacidini.Item Revision and phylogeny of the caddisfly subfamily Protoptilinae (Trichoptera: Glossosomatidae) inferred from adult morphology and mitochondrial DNA(Magnolia Press, 2013) Robertson, Desiree R.; Holzenthal, Ralph W.Protoptilinae Ross, 1956, is the most diverse subfamily belonging to the saddle- or tortoise-case-making caddisfly family Glossosomatidae Wallengren, 1891. The subfamily has a disjunct distribution: 5 genera are known from the East Palaearctic and Oriental regions; the remaining 13 are restricted to the Nearctic and Neotropical regions. Monophyly of Protoptilinae and each of 17 genera was tested using 80 taxa, 99 morphological characters, and mitochondrial DNA (COI). Additionally, homologies of morphological characters were assessed across genera and a standardized terminology for those structures was established. Mitochondrial DNA data were unavailable for 55 of the 80 taxa included in this study. To test the effects of the missing molecular data, 5 different datasets were analyzed using both parsimony and Bayesian methods. There was incongruence between the COI and morphological data, but results suggest the inclusion of COI data in a combined analysis, although incomplete, improved the overall phylogenetic signal. Bayesian and parsimony analyses of all 5 datasets strongly supported the monophyly of Protoptilinae. Monophyly of the following genera was also supported: Canoptila Mosely, 1939; Culoptila Mosely, 1954; Itauara Müller, 1888; Mastigoptila Flint, 1967; Mortoniella Ulmer, 1906; Protoptila Banks, 1904; and Tolhuaca Schmid, 1964. Several taxonomic changes were necessary for classification to reflect phylogeny accurately. Accordingly, Matrioptila Ross, 1938; Poeciloptila Schmid, 1991; Temburongpsyche Malicky, 1992; and Nepaloptila Kimmins, 1964, are designated new junior synonyms of Padunia Martynov, 1910. Additionally, the endemic Caribbean genera Campsiophora Flint, 1964, and Cubanoptila Sykora, 1973, are designated new junior synonyms of Cariboptila Flint, 1964. Diagnoses and a key to the subfamilies of Glossosomatidae and world genera of Protoptilinae incorporating these taxonomic changes are provided.Item Revision of the Neotropical caddisfly genus Itauara Müller, 1888 (Trichoptera, Glossosomatidae)(Pensoft Publishers, 2011) Robertson, Desiree R.; Holzenthal, Ralph W.Systematics of the genus Itauara Müller, 1888 are reviewed. A generic diagnosis, illustrations, and descriptions are provided for males. The genus can be identified by several features of the male genitalia including an extremely reduced phallobase and a phallic apparatus that consists of a sclerotized dorsal sheath covering a very membranous ventral portion. A total 18 species are described as new: Itauara alexanderi, sp. n. (Brazil), I. bidentata, sp. n. (Guyana), I. blahniki, sp. n. (Brazil), I. charlotta, sp. n. (Brazil), I. emilia, sp. n. (Brazil), I. flinti, sp. n. (Brazil), I. guyanensis, sp. n. (Guyana), I. jamesii, sp. n. (Brazil), I. julia, sp. n. (Brazil), I. lucinda, sp. n. (Brazil), I. ovis, sp. n. (Guyana, Venezuela), I. peruensis, sp. n. (Peru), I. rodmani, sp. n. (Brazil), I. simplex, sp. n. (Brazil), I. spiralis, sp. n. (Guyana), I. stella, sp. n. (Brazil), I. tusci, sp n. (Brazil), and I. unidentata, sp. n. (Guyana). These additions bring the total fauna of Itauara to 22 species.Item Two new species and a new record of Protoptila from Bolivia (Trichoptera: Glossosomatidae: Protoptilinae)(Oxford University Press, 2008) Robertson, Desiree R.; Holzenthal, Ralph W.The caddisfly genus Protoptila (Trichoptera: Glossosomatidae: Protoptilinae) has never before been recorded from Bolivia. Protoptila dubitans Mosely 1939 is now reported from Bolivia for the first time. Additionally, two new species from Bolivia, P. diablita and P. julieta are described. Protoptila julieta is most closely related to P. misionensis Flint 1972. Protoptila diablita is the first known species in the genus to possess scale-like setae. These scales are hypothesized to aid in the dispersal of male sex pheromones.