Browsing by Author "Blahnik, Roger J."
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Item The caddisfly genus Protoptila in Costa Rica (Trichoptera: Glossosomatidae)(Magnolia Press, 2006) Holzenthal, Ralph W.; Blahnik, Roger J.Nineteen species of Protoptila known from Costa Rica are revised to include 11 previously described species (P. bicornuta Flint 1963, P. boruca Flint 1974, P. burica Flint 1974, P. cana Flint 1974, P. ixtala Mosely 1937, P. laterospina Flint 1967, P. orotina orotina Flint 1974, P. spirifera Flint 1974, P. talamanca Flint 1974, P. tica Bueno-Soria 1984, and P. tojana Mosely 1954) and 8 new species (P. altura, P. bribri, P. chitaria, P. cristula, P. kjeri, P. jolandae, P. strepsicera, and P. trichoglossa). Illustrations, diagnoses, descriptions, and distribution records are provided for all species.Item Checklist of the Trichoptera (Insecta) of Brazil I(2004) Paprocki, Henrique; Holzenthal, Ralph W.; Blahnik, Roger J.We present here a list of Trichoptera species recorded in the literature from Brazil as of September 2003. The total number of species recorded for Brazil is 378. The most diverse family is Hydropsychidae with 103 species; the second most diverse is Hydroptilidae with 50 species, followed by Leptoceridae and Philopotamidae with 41 each. Distributions by state and additional literature relevant to Brazilian Trichoptera are also presented.Item Chimarra of Sabah and Sarawak, northern Borneo (Trichoptera: Philopotamidae)(Brill Academic Publishers, 2009) Blahnik, Roger J.; Holzenthal, Ralph W.; Huisman, JolandaThirty new species of Chimarra Stephens (Trichoptera: Philopotamidae) from the Malaysian part of Borneo (Sabah and Sarawak) are described, and eight previously described species are redescribed (Chimarra dulitensis Kimmins, 1955 and C. demeter Malicky, 2000, originally described from the Malaysian part of Borneo; and also C. phlegyas Malicky, 2008; C. polyneikes Malicky, 2008; C. prokrustes Malicky, 2008; C. terramater Malicky, 2008; C. thaumas Malicky, 2008; and C. tityos Malicky, 2008, originally described from the Indonesian part of Borneo, but now also recorded from the Malaysian part of Borneo). New species described include: Chimarra antheae, C. caduca, C. chanchuluni, C. cuspidata, C. cygnus, C. danumensis, C. dejongi, C. denticula, C. devogeli, C. drepane, C. fuilianae, C. gyrospina, C. jannekae, C. karlijnae, C. kinabaluensis, C. lambi, C. liwaguensis, C. noloyan, C. noohi, C. phillipsae, C. physanoton, C. preapicalis, C. scolops, C. silausilau, C. sinitorum, C. stenodactylus, C. vantoli, C. vanwelzeni, C. ventritropis, and C. xiphosella. Two major groups, the Chimarra tsudai group and the Chimarra digitata group are recognized, based on characters of the male genitalia and wing venation, to include most of these species.Item DNA barcode data confirm new species and reveal cryptic diversity in Chilean Smicridea (Smicridea) (Trichoptera:Hydropsychidae)(The North American Benthological Society, 2010) Pauls, Steffen U.; Blahnik, Roger J.; Zhou, Xin; Wardwell, C. Taylor; Holzenthal, Ralph W.Mitochondrial deoxyribonucleic acid (mtDNA) sequence data have been both heralded and scrutinized for their ability or lack thereof to discriminate among species for identification (DNA barcoding) or description (DNA taxonomy). Few studies have systematically examined the ability of mtDNA from the DNA barcode region (658 base pair fragment of the 59 terminus of the mitochondrial cytochrome c oxidase I gene) to distinguish species based on range-wide sampling of specimens from closely related species. Here we examined the utility of DNA barcode data for delimiting species, associating life stages, and as a potential genetic marker for phylogeographic studies by analyzing a rangewide sample of closely related Chilean representatives of the caddisfly genus Smicridea subgenus Smicridea. Our data revealed the existence of 7 deeply diverged, previously unrecognized lineages and confirmed the existence of 2 new species: Smicridea (S.) patinae, new species and Smicridea (S.) lourditae, new species. Based on our current taxonomic evaluation, we considered the other 5 lineages to be cryptic species. The DNA barcode data proved useful in delimiting species within Chilean Smicridea (Smicridea) and were suitable for life-stage associations. The data also contained sufficient intraspecific variation to make the DNA barcode a candidate locus for widespread application in phylogeographic studies.Item New distribution and species records of Trichoptera from Southern and Southeastern Brazil(2004) Blahnik, Roger J.; Paprocki, Henrique; Holzenthal, Ralph W.We provide here a list of new species records of Trichoptera (Insecta) collected in Southern and Southeastern Brazil. We report 21 new distribution records for the country of Brazil. We also provide new distribution records for 92 species of Trichoptera for the states of Minas Gerais, São Paulo, Rio de Janeiro, Paraná and Santa Catarina.Item New Neotropical species of Chimarra (Trichoptera, Philopotamidae)(Pensoft Publishers, 2012) Blahnik, Roger J.; Holzenthal, Ralph W.Ten new Neotropical species of Chimarra are described in the subgenera Chimarra, Chimarrita, and Otarrha. New species in the subgenus Chimarra include, in the C. ortiziana group: C. calori sp. n. (southeastern Brazil) and C. onchyrhina sp. n. (Venezuela); in the C. picea group: C. inchoata sp. n. (Venezuela), C. nicehuh sp. n. (Venezuela), and C. sunima sp. n. (Colombia); and in the C. poolei group: C. cauca sp. n. (Colombia) and C. desirae sp. n. (Bolivia). New species in the subgenus Chimarrita include, in the C. simpliciforma group: C. curvipenis sp. n. (SE Brazil) and C. latiforceps sp. n. (SE Brazil). A single new species in the subgenus Otarrha is also described: C. soroa sp. n. (Cuba). Males and females for all of the new species are illustrated, except for C. desirae, for which female specimens were unavailable. Additionally, the female of C. (Chimarrita) camella, which was previously unknown, is illustrated.Item New species of Smicridea (Rhyacophylax) (Trichopera: Hydropsychidae) from Costa Rica(American Entomological Society, 1995) Holzentahl, Ralph W.; Blahnik, Roger J.Three new species in lhe caddjsfly genus Smicridea, subgenus Rhyacophylax, are described from Costa Rica and 25 additional species in the two recognized subgenera are recorded from the country. Illustrations of the male and female genitalia and wing coloration are provided for each of the new species along with a map of their distributions. Two of the new species, S. singri and S. nemorosa, are in the signota species group. and the third, S. rapanti, is a member of the peruana species group.Item Order Trichoptera Kirby, 1813 (Insecta), Caddisflies(Magnolia Press, 2007) Holzenthal, Ralph W.; Blahnik, Roger J.; Prather, Aysha L.; Kjer, Karl M.The taxonomy, diversity, and distribution of the aquatic insect order Trichoptera, caddisflies, are reviewed. The order is among the most important and diverse of all aquatic taxa. Larvae are vital participants in aquatic food webs and their presence and relative abundance are used in the biological assessment and monitoring of water quality. The species described by Linnaeus are listed. The morphology of all life history stages (adults, larvae, and pupae) is diagnosed and major features of the anatomy are illustrated. Major components of life history and biology are summarized. A discussion of phylogenetic studies within the order is presented, including higher classification of the suborders and superfamilies, based on recent literature. Synopses of each of 45 families are presented, including the taxonomic history of the family, a list of all known genera in each family, their general distribution and relative species diversity, and a short overview of family-level biological features. The order contains 600 genera, and approximately 13,000 species.Item Phylogeny of Trichoptera.(2003) Kjer, Karl M.; Holzenthal, Ralph W.; Blahnik, Roger J.Item Review and redescription of species in the Oecetis avara group, with the description of 15 new species (Trichoptera, Leptoceridae)(Pensoft Publishers, 2014) Blahnik, Roger J.; Holzenthal, Ralph W.The O. avara group of Oecetis is formally defined to include 4 described species, O. avara (Banks), O. disjuncta (Banks), O. elata Denning & Sykora, and O. metlacenis Bueno-Soria, and 15 new species. Oecetis marquesi Bueno-Soria, previously considered a member of the O. avara group, is treated as incertae sedis to species group, but is also redescribed and treated in the current work. New species described here (with their respective distributions) include: O. acciptrina (Costa Rica, Panama, Ecuador), O. agosta (Mexico), O. angularis (Guatemala to Ecuador), O. apache (SW USA), O. campana (Ecuador), O. constricta (Mexico to Ecuador, Venezuela, and Trinidad), O. houghtoni (North America), O. maritza (Costa Rica), O. mexicana (Mexico to Ecuador), O. patula (Guatemala, Nicaragua), O. protrusa (Mexico to Ecuador), O. sordida (Mexico, USA, Canada), O. tumida (Costa Rica), O. uncata (Costa Rica), and O. verrucula (Mexico to Costa Rica). A key to the species is also provided.Item Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae)(Magnolia Press, 2011) Blahnik, Roger J.; Holzenthal, Ralph W.Species of Mortoniella are revised for the southern part of the South American continent, including the countries of Brazil, Paraguay, Uruguay, Argentina and Chile. The M. velasquezi species group is diagnosed for the first time and the M. bilineata species group is more critically defined. Males of all species from the region, except M. armata (Jacquemart, 1963), are figured, and a key is provided for their identification. Included in the revision are 11 described species and 22 new species. Previously described species that are redescribed and figured include the following: Mortoniella albolineata Ulmer, 1907; M. argentinica Flint, 1974; M. catarinensis (Flint, 1974); M. collegarum (Rueda & Gibon, 2008); M. guairica (Flint, 1974); M. ormina (Mosely, 1939); M. pocita (Flint, 1983); M. punensis (Flint, 1983); M. teutona (Mosely, 1939) M. unota (Mosely, 1939); and M. wygodzinskii (Schmid, 1958). Mortoniella collegarum, originally described from Bolivia and Argentina, is here reported from Chile, representing the first distributional record of Mortoniella for that country. It is assigned to the M. ormina species group. On morphological considerations, M. argentinica is removed from the M. bilineata species group and left as a species incertae sedis as to species group. Based on its overall similarity to M. argentinica, M. spinulata (Flint, 1991), from Colombia, is removed from the M. leroda species group and left as a species incertae sedis as to species group. New species of Mortoniella described here, followed by their respective areas of distribution, include, for the M. leroda species group: Mortoniella acauda (SE Brazil), M. agosta (SE Brazil), M. asym- metris (Paraguay), M. crescentis (SE Brazil), M. dolonis (SE Brazil), M. guahybae (SE Brazil), M. hystricosa (SE Brazil), M. intervales (SE Brazil), M. latispina (SE Brazil), M. longispina (SE Brazil), M. paraguaiensis (Paraguay), M. parauna (SE Brazil), M. paraunota (Argentina, SE Brazil), M. pumila (SE Brazil), M. pusilla (SE Brazil), M. truncata (SE Brazil), and M. uruguaiensis (SE Brazil, Uruguay); for the M. ormina species group: M. alicula (SE Brazil); and for the M. velasquezi species group: M. bocaina (SE Brazil), M. froehlichi (SE Brazil), and M. tripuiensis (SE Brazil). A new species unplaced as to species group is M. meloi (SE Brazil).Item Revision of the genus Culoptila (Trichoptera: Glossosomatidae)(Magnolia Press, 2006) Blahnik, Roger J.; Holzenthal, Ralph W.The caddisfly genus Culoptila is revised to include 17 previously described species and 9 new species. Illustrations of the male genitalia and a key are provided for all species. In addition, illustrations of male head and thoracic features, larval characters and cases, and female genitalia are included. New species described here include: Culoptila bidentata (Costa Rica), C. buenoi (Mexico), C. cascada (Costa Rica), C. hamata (Costa Rica), C. pararusia (Mexico), C. plummerensis (eastern United States), C. tapanti (Costa Rica), C. unispina (Costa Rica, Panama), and C. vexillifera (Guatemala).Item Revision of the Mexican and Central American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae)(Magnolia Press, 2008) Blahnik, Roger J.; Holzenthal, Ralph W.The genus Mexitrichia Mosely, 1937 (27 described species) is synonymized with Mortoniella Ulmer, 1906 (22 described species) and a revised generic description is provided for the genus. These species are placed in 4 recognized species groups. Mexican and Central American species of Mortoniella are revised to include 6 species formerly placed in Mex- itrichia and 22 new species from Costa Rica, Panama, and Mexico. New species combinations for these regional species include: Mortoniella florica (Flint, 1974), M. leroda (Mosely, 1937), M. meralda (Mosely, 1954), M. pacuara (Flint, 1974), M. rancura (Mosely, 1954), and M. rovira (Flint, 1974). New species of Mortoniella described here (followed by the country of provenance) include M. akantha (Costa Rica) M. anakantha (Costa Rica) M. aviceps (Costa Rica, Pan- ama) M. brachyrhachos (Mexico), M. buenoi (Mexico), M. carinula (Costa Rica), M. caudicula (Costa Rica), M. falcic- ula (Mexico), M. mexicana (Mexico) M. munozi (Costa Rica, Panama), M. opinionis (Costa Rica), M. panamensis (Panama), M. papillata (Costa Rica), M. pectinella (Panama), M. propinqua (Costa Rica), M. redunca (Costa Rica), M. rodmani (Costa Rica), M. sicula (Costa Rica), M. stilula (Costa Rica), M. tapanti (Costa Rica, Panama), M. taurina (Costa Rica, Panama), and M. umbonata (Panama). Males of all known Mexican and Central American species are illus- trated or reillustrated and a key is provided for males of the species.Item Systematics of the Neotropical caddisfly genus Notidobiella Schmid (Trichoptera, Sericostomatidae), with the description of 3 new species(Pensoft Publishers, 2010) Holzenthal, Ralph W.; Blahnik, Roger J.Three new species of Notidobiella Schmid (Insecta: Trichoptera) are described from South America: Notidobiella amazoniana sp. n. (Brazil), N. brasiliana sp. n. (Brazil), and N. ecuadorensis sp. n. (Ecuador). In addition, the 3 previously described species in the genus, N. chacayana Schmid, N. inermis Flint, and N. parallelipipeda Schmid, all endemic to southern Chile, are redescribed and illustrated, including the females of each species for the first time, and a key to males of the species in the genus is provided. The occurrence of Notidobiella in Brazil and Ecuador represents a signifi cant extension of the range of the genus beyond southern Chile where it previously was thought to be endemic. The biogeography of Sericostomatidae and other austral South American Trichoptera is reviewed. The presence of the family in South America may not be part of a “transantarctic” exchange, but instead may represent an earlier occurence in the region. The distribution of Notidobiella in tropical South America likely represents recent dispersal from southern South America to the north.Item Three new species of Helicopsyche von Siebold (Trichoptera: Helicopsychidae) from Brazil(Magnolia Press, 2016) Holzenthal, Ralph W.; Blahnik, Roger J.; Calor, Adolfo R.Three new species of Helicopsyche subgenus Feropsyche (Trichoptera: Helicopsychidae) from southern and southeastern Brazil are diagnosed, described, and illustrated: Helicopsyche angeloi sp. nov. (Holotype male deposited in MZSP: Minas Gerais, Rio de Janeiro and São Paulo States), H. guara sp. nov. (Holotype male deposited in MZSP: Minas Gerais and Santa Catarina States) and H. lazzariae sp. nov. (Holotype male in MZSP: Paraná State). The first two species are similar to H. cipoensis Johanson & Malm, 2006, and H. timbira Silva, Santos & Nessimian, 2014, in having similarly shaped inferior appendages. However, the new species differs in the morphology and distribution of setae on the inferior appendages and tergum X. Helicopsyche lazzariae sp. nov. is unlike any other described species in the subgenus Feropsyche because of its broad deltoid-shaped inferior appendage and absence of a basomesal process on the inferior appendage. Characters important for diagnosing and describing new species of Helicopsyche (Feropsyche) are discussed. Finally, an updated checklist of the 23 species recorded for Brazil is presented.Item The Trichoptera barcode initiative: a strategy for generating a species-level Tree of Life(The Royal Society Publishing, 2016) Zhou, Xin; Frandsen, Paul B.; Holzenthal, Ralph W.; Beet, Clare R.; Bennett, Kristi R.; Blahnik, Roger J.; Bonada, Nu´ria; Cartwright, David; Chuluunbat, Suvdtsetseg; Cocks, Graeme V.; Collins, Gemma E.; deWaard, Jeremy; Dean, John; Flint, Oliver S. Jr; Hausmann, Axel; Hendrich, Lars; Hess, Monika; Hogg, Ian D.; Kondratieff, Boris C.; Malicky, Hans; Milton, Megan A.; Morinie`re, Je´roˆme; Morse, John C.; Ngera Mwangi, Francois; Pauls, Steffen U.; Razo Gonzalez, Marı´a; Rinne, Aki; Robinson, Jason L.; Salokannel, Juha; Shackleton, Michael; Smith, Brian; Stamatakis, Alexandros; StClair, Ros; Thomas, Jessica A.; Zamora-Munoz, Carmen; Ziesmann, Tanja; Kjer, Karl M.DNA barcoding was intended as a means to provide species-level identifications through associating DNA sequences from unknown specimens to those from curated reference specimens. Although barcodes were not designed for phylogenetics, they can be beneficial to the completion of the Tree of Life. The barcode database for Trichoptera is relatively comprehensive, with data from every family, approximately two-thirds of the genera, and one-third of the described species. Most Trichoptera, as with most of life’s species, have never been subjected to any formal phylogenetic analysis. Here, we present a phylogeny with over 16 000 unique haplotypes as a working hypothesis that can be updated as our estimates improve. We suggest a strategy of implementing constrained tree searches, which allow larger datasets to dictate the backbone phylogeny, while the barcode data fill out the tips of the tree. We also discuss how this phylogeny could be used to focus taxonomic attention on ambiguous species boundaries and hidden biodiversity. We suggest that systematists continue to differentiate between ‘Barcode Index Numbers’ (BINs) and ‘species’ that have been formally described. Each has utility, but they are not synonyms. We highlight examples of integrative taxonomy, using both barcodes and morphology for species description. This article is part of the themed issue ‘From DNA barcodes to biomes’.